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Cited article:

Phosphorylation of Ser-80 of VP1 and Ser-254 of VP2 is essential for human BK virus propagation in tissue culture

Pei-Lain Chen, Pang-Hung Hsu, Chiung-Yao Fang, et al.
Journal of General Virology 92 (11) 2637 (2011)
https://doi.org/10.1099/vir.0.033282-0