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Cited article:

Repeated exposures to UVB induce differentiation rather than senescence of human keratinocytes lacking p16INK-4A

Véronique Bertrand-Vallery, Emmanuelle Boilan, Noëlle Ninane, et al.
Biogerontology 11 (2) 167 (2010)
https://doi.org/10.1007/s10522-009-9238-y