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Cited article:

The Common γ-Chain Cytokines IL-2, IL-7, IL-15, and IL-21 Induce the Expression of Programmed Death-1 and Its Ligands

Audrey L. Kinter, Emily J. Godbout, Jonathan P. McNally, et al.
The Journal of Immunology 181 (10) 6738 (2008)
https://doi.org/10.4049/jimmunol.181.10.6738